A new species of Patagonian megaraptor has been described.  Joaquinraptor casali provides further insight into this enigmatic Cretaceous dinosaur family.  The fossil material casts light on these giant-clawed theropods, as this is only the second member of the Megaraptoridae family to have been discovered with extensive skull remains.  The study has been published in the journal “Nature Communications”.

Intriguingly, the field team found a crocodyliform humerus between the lower jaw bones of Joaquinraptor.  This suggests that that the new megaraptor may have been eating the crocodyliform when it died.  However, this is only an assertion, it is not definitive.  The limb bone could have ended up lodged between the dentaries as a result of being washed into the submerged mouth of the dinosaur’s carcase.

Joaquinraptor casali life reconstruction. The new megaraptor dinosaur species Joaquinraptor casali in what is now central Patagonia, Argentina roughly 67 million years ago. The carnivore holds a crocodyliform (= extinct crocodile relative) front leg in its mouth, in keeping with the discovery of a crocodyliform forelimb bone between the lower jaws. Picture credit: Andrew McAfee, Carnegie Museum of Natural History.

Picture credit: Andrew McAfee, Carnegie Museum of Natural History.

The upper arm bone of the crocodyliform bears tooth marks and the bone was touching some of the megaraptor’s teeth.  So, some kind of interspecific interaction seems likely. Perhaps, the theropod and the crocodyliform were fighting.  For example, the two predators could have been battling over a food resource.

Possible evidence of diet or predation in the new megaraptor dinosaur species Joaquinraptor casali. (a) Dentaries (= tooth-bearing lower jaw bones) of Joaquinraptor and an associated crocodyliform (= extinct crocodile relative) forelimb bone in the position in which they were found, prior to their separation in the laboratory. Detail (b) view showing a portion of the croc forelimb bone in contact with the teeth of Joaquinraptor. The croc bone (c) after its separation from the Joaquinraptor dentaries. Picture credit: Lucio M. Ibiricu et al.

Picture credit: Lucio M. Ibiricu et al

Joaquinraptor casali

Joaquinraptor seems to be quite typical for a megaraptor.  It had a long, low skull lined with serrated teeth and powerful forelimbs. The two innermost talons of the hand were tipped with large claws.  The genus name honours Joaquín, the son of the first author (Lucio M. Ibiricu) and the informal name given to the locality when the skeleton of the taxon was discovered (Valle Joaquín). Dr Gabriel Andrés Casal inspired the species epithet.  Dr Casal has made a significant contribution to our understanding of Cretaceous dinosaurs from Argentina.  In addition, he was responsible for the formal recognition and naming of the formation from which this megaraptorid was recovered (the Lago Colhué Huapi Formation).

A skeletal reconstruction of the new megaraptor dinosaur species Joaquinraptor casali showing the bones preserved (in blue). Other parts of the skeleton and body outline silhouette are based on closely related species. Photos of many of the preserved bones in various views surround the skeletal reconstruction. Skeletal reconstruction by T.K. Robinson and Andrew McAfee. Picture credit: Lucio M. Ibiricu et al.

Picture credit: Lucio M. Ibiricu et al

The Geologically Youngest Member of the Megaraptoridae

The researchers estimate that Joaquinraptor was around seven metres in length.  It probably weighed a tonne. This newly described theropod is also likely to be the geologically youngest megaraptor ever discovered.  Megaraptors survived until the end of the Cretaceous, 66 million years ago.  The partial, skeleton comes from the Chubut Province (southern Patagonia).  It was studied by Dr Lucio Ibiricu of Argentina’s Instituto Patagónico de Geología y Paleontología, and several other scientists including Dr Matt Lamanna of the Carnegie Museum of Natural History.

Dr Lamanna commented:

“At the Carnegie Museum of Natural History, we’re always seeking to better comprehend the history of life on our planet.  Every year, our team of experts conducts field work all over the world, helping to ensure that we’re on the cutting edge of research.  New revelatory dinosaur discoveries, like Joaquinraptor casali, deepen our understanding of how extraordinary creatures lived and advance understanding of the natural world, both past and present.”

Fossils of the new megaraptor dinosaur species J. casali in the quarry, laying in the positions they had laid for approximately 67 million years. A dentary, teeth, ribs, and several other bones are visible. Picture credit: Marcelo Luna, Laboratorio de Paleontología de Vertebrados Dr. Rubén Martínez, Universidad Nacional de la Patagonia San Juan Bosco.

Picture credit: Everything Dinosaur

An analysis of a cross section of the tibia revealed that this dinosaur was at least nineteen years old when it perished. However, whilst the researchers considered the specimen to have been sexually mature, it was not fully grown.

Everything Dinosaur acknowledges the assistance of a media release from the Carnegie Museum of Natural History in the compilation of this article.

The scientific paper: “Latest Cretaceous megaraptorid theropod dinosaur sheds light on megaraptoran evolution and palaeobiology” by Lucio M. Ibiricu, Matthew C. Lamanna, Bruno N. Alvarez, Ignacio A. Cerda, Julieta L. Caglianone, Noelia V. Cardozo, Marcelo Luna and Rubén D. Martínez published in Nature Communications.

The Everything Dinosaur website: Dinosaur Models.

The thick-skulled, dome-headed pachycephalosaurs are among the most enigmatic and yet, poorly known of all the Dinosauria.  For example, these ornithischians, with their elaborate cranial ornamentation are thought to have evolved complex social behaviours.  After all, the idea that those super thick skulls played a role in butting contests has been around for a long time.  The pachycephalosaurs have a new member.  The recently described Zavacephale rinpoche represents the oldest pachycephalosaur known to science.  It roamed Mongolia around 115 to 108 million years ago.  Its discovery helps palaeontologists to reconstruct the origins and the early evolution of the pachycephalosaurians.

The fossil represents a sub-adult, but the skull already has the distinctive dome. This indicates that pachycephalosaurs developed features relating to combat or display before they reached adulthood. Picture credit: Masato Hattori.

Picture credit: Masato Hattori

Zavacephale rinpoche

The fossil specimen, thought to represent a sub-adult, was discovered in the Khuren Dukh locality of the Eastern Gobi Basin. Not only is the specimen the oldest pachycephalosaur known to science, the skeleton is the most complete for this type of dinosaur found to date.

Corresponding author for the study, Lindsay Zanno, an associate research professor at North Carolina State University commented:

“Pachycephalosaurs are iconic dinosaurs, but they’re also rare and mysterious.”

The fossil material was found by Tsogtbaatar Chinzorig (Mongolian Academy of Sciences), the lead author of the study published in the journal “Nature”. The Z. rinpoche specimen was not fully grown when it died. However, it already bore a fully formed dome, though without much of the additional ornamentation found on other pachycephalosaur fossils.  The genus name is derived from zava which means “root” or “origin” in Tibetan and “cephal” which translates from Latin meaning head. The species name is derived from the Tibetan phrase for “precious one” as the domed skull was exposed on a cliff like a cabochon jewel.

Lindsay Zanno of the Department of Biological Sciences, (North Carolina State University), highlighting the morphology of the Zavacephale rinpoche skull. Picture credit: Alfio Alessandro Chiarenza.

Picture credit: Alfio Alessandro Chiarenza

Predating All Other Pachycephalosaur Fossils by at Least 14 Million Years

Zavacephale rinpoche pushes back fossil evidence of the frontoparietal dome by at least fourteen million years.  Furthermore, the skeleton, which is more than fifty percent complete, preserves regions of the body not scientifically described in any other pachycephalosaur.  For example, scientists could examine an almost complete tail covered in ossified tendons.  In addition, bones from the hand, a first for a pachycephalosaur, were found.

Intriguingly, the specimen had a gastrolith mass preserved in the stomach region.  These stomach stones suggest an omnivorous diet.  The stones ground up tough plant food to help with digestion.  It had been thought that these dinosaurs were herbivores.  However, recent discoveries such as fang-like teeth had led some researchers to speculate that they were carnivorous ornithischians.  The discovery of these stomach stones lends support to the theory that pachycephalosaurs probably ate both plants and animals, that they were omnivores.

To read a blog post from Everything Dinosaur: Pachycephalosaurus – Was it Carnivorous?

Highlighting the significance of Zavacephale rinpoche Tsogtbaatar Chinzorig stated:

“Z. rinpoche predates all known pachycephalosaur fossils to date by about 15 million years. It was a small animal – about three feet or less than one metre long, and the most skeletally complete specimen yet found.”

Chinzorig added:

“Z. rinpoche is an important specimen for understanding the cranial dome development of pachycephalosaurs, which has been debated for a long time due to the absence of early diverging or pre-Late Cretaceous species and the fragmentary nature of nearly all pachycephalosaurian fossils.”

Calculating the Age of a Dinosaur

A thin slice was carefully removed from a lower leg bone.  This slice was then subjected to histological analysis to calculate the dinosaur’s age when it died.  For the first time, palaeontologists had an almost complete pachycephalosaur skull with associated limb bones to help calculate the growth stage of the skeleton.

Lindsay Zanno explained:

“Pachycephalosaurs are all about the bling, but we can’t use flashy signalling structures alone to figure out what species they belong to or what growth stage they’re in because some cranial ornamentation changes as animals mature. We age dinosaurs by looking at growth rings in bones, but most pachycephalosaur skeletons are just isolated, fragmentary skulls. Z. rinpoche is a spectacular find because it has limbs and a complete skull, allowing us to couple growth stage and dome development for the first time.”

The fossil material has pushed back the fossil record of pachycephalosaurs by at least fourteen million years.  In addition, the research team have a much more complete fossil specimen to study.  Moreover, Zavacephale rinpoche demonstrates that dome heads developed in pachycephalosaurs before they reached adulthood.

Lindsay Zanno quipped:

“If you need to headbutt yourself into a relationship, it’s a good idea to start rehearsing early.”

Late Cretaceous Pachycephalosaurs

Most pachycephalosaur fossils are found in Upper Cretaceous deposits, primarily deposits laid down in the Campanian and Maastrichtian.

Austin the Pachycephalosaurus (P. wyomingensis).

Most specimens have been described based on cranial material.  These dinosaurs were bipedal.  Pachycephalosaurus had five fingers on each hand.  The image above of P. wyomingensis shows the PNSO model – Austin the Pachycephalosaurus.  This figure has been widely praised for its accuracy.

To view the range of PNSO models and figures in stock: PNSO Age of Dinosaurs Models.

An Important Discovery

The researchers stressed the significance of this fossil discovery.  Lindsay Zanno stated:

“This specimen is a once-in-a-lifetime discovery. It is remarkable for being the oldest definitive pachycephalosaur, pushing back the fossil record of this group by at least 15 million years, but also because of how complete and well-preserved it is. Z. rinpoche gives us an unprecedented glimpse into the anatomy and biology of pachycephalosaurs, including what their hands looked like and that they used stomach stones to grind food.”

Chinzorig added:

“The newly recovered materials of Z. rinpoche, such as the hand elements, the stomach stones (gastroliths), and an articulated tail with covered tendons, reshape our understanding of the paleobiology, locomotion, and body plan of these ‘mysterious’ dinosaurs.”

Everything Dinosaur acknowledges the assistance of a media release from the North Carolina State University in the compilation of this article.

The scientific paper: “A domed pachycephalosaur from the early Cretaceous of Mongolia” by Tsogtbaatar Chinzorig, Ryuji Takasaki, Junki Yoshida, Ryan T. Tucker, Batsaikhan Buyantegsh, Buuvei Mainbayar, Khishigjav Tsogtbaatar and Lindsay E. Zanno published in the journal Nature.

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